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Chinese Acad Sci, Inst Appl Ecol, CAS Key Lab Forest Ecol & Management, Shenyang 110016, Peoples R China
Northeast Normal Univ, Sch Geog Sci, Key Lab Geog Proc & Ecol Secur Changbai Mt, Minist Educ, Changchun 130024, Peoples R China Chinese Acad Sci, Inst Appl Ecol, CAS Key Lab Forest Ecol & Management, Shenyang 110016, Peoples R China