[1] Hokkaido University School of Medicine,Department of Ophthalmology
[2] Tokyo Metropolitan Institute for Neuroscience,Department of Molecular Neurobiology
[3] Hokkaido University School of Medicine,Department of Biochemistry
[4] Nippon Medical College,Department of Anatomy
[5] Kyushu University,Department of Molecular and Cellular Biology, Medical Institute of Bioregulation
[6] Tohoku University Graduate School of Medicine,Division of Developmental Genetics, Center for Translational and Advanced Animal Research on Human Diseases
[7] Hokkaido University School of Medicine,Department of Ophthalmology