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Univ Edinburgh, Sch Biomed Sci, Ctr Discovery Brain Sci, Hugh Robson Bldg,15 George Sq, Edinburgh EH8 9DX, Midlothian, Scotland Univ Edinburgh, Sch Biomed Sci, Ctr Discovery Brain Sci, Hugh Robson Bldg,15 George Sq, Edinburgh EH8 9DX, Midlothian, Scotland