[1] Imperial Coll London, Dept Life Sci, London, England
[2] Macquarie Univ, Dept Biol Sci, N Ryde, NSW, Australia
[3] Tsinghua Univ, Dept Earth Syst Sci, Minist Educ, Key Lab Earth Syst Modelling, Beijing, Peoples R China
[4] Univ New South Wales, Climate Change Res Ctr ESSRC, Sydney, NSW, Australia
[5] Univ Reading, Dept Geog & Environm Sci, Reading, Berks, England