[1] Chinese Academy of Sciences,Key Laboratory of Alpine Ecology and Biodiversity, Institute of Tibetan Plateau Research
[2] University of Chinese Academy of Sciences,Département des Sciences Fondamentales
[3] Université du Québec à Chicoutimi,Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems, Provincial Key Laboratory of Applied Botany, South China Botanical Garden
[4] Chinese Academy of Sciences,undefined
[5] CAS Center for Excellence in Tibetan Plateau Earth Sciences,undefined
[6] Instituto Pirenaico de Ecología (IPE-CSIC),undefined