[1] University of the Chinese Academy of Sciences,College of Life Sciences
[2] Chinese Academy of Sciences,Key Laboratory of Alpine Ecology, Institute of Tibetan Plateau Research
[3] Chinese Academy of Sciences,Key Laboratory of Ecosystem Network Observation and Modeling, Institute of Geographic Sciences and Natural Resources Research
[4] University of Wisconsin-Madison,Department of Soil Science
[5] Chinese Academy of Sciences,Northwest Institute of Plateau Biology, Key Laboratory of Adaptation and Evolution of Plateau Biota