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Huazhong Cent China Agr Univ, Coll Anim Sci & Technol,Minist Educ, Key Lab Swine Genet & Breeding,Minist Agr, Key Lab Agr Anim Genet Breeding & Reprod, Wuhan, Hubei, Peoples R China Huazhong Cent China Agr Univ, Coll Anim Sci & Technol,Minist Educ, Key Lab Swine Genet & Breeding,Minist Agr, Key Lab Agr Anim Genet Breeding & Reprod, Wuhan, Hubei, Peoples R China