[1] University of Guelph,Department of Integrative Biology
[2] University of Toronto – Mississauga,Department of Ecology and Evolutionary Biology
[3] University of Colorado Boulder,Institute of Arctic and Alpine Research, Department of Ecology and Evolutionary Biology
[4] Northern Arizona University,Center for Ecosystem Science and Society and Department of Biological Sciences
[5] Michigan Technological University,College of Forest Resources and Environmental Science
[6] USDA Forest Service Northern Research Station,undefined