[1] Kyoto Univ, Wildlife Res Ctr, Sakyo Ku, 2-24 Tanaka Sekiden Cho, Kyoto 6068203, Japan
[2] FFPRI, 1 Matsunosato, Tsukuba, Ibaraki 3058687, Japan
[3] Amami Wild Anim Res Ctr, 2662 Ogachi,Tatsugo Cho, Kagoshima 8940105, Japan
[4] Amami Wildlife Res Ctr Co Ltd, 10-11-2F Naze Suehiro Cho, Kagoshima 8940027, Japan