[1] Univ Lyon 1, CNRS, UMR 5558, Lab Biometrie & Biol Evolut, F-69622 Villeurbanne, France
[2] SISSA, I-34014 Trieste, Italy
[3] Ist Nazl Fis Nucl, Sez Trieste, Trieste, Italy
[4] Politecn Torino, Turin, Italy
[5] Univ Washington, IHME, Seattle, WA 98195 USA