[1] Univ Hamburg, Bioctr Grindel, Inst Zool, D-20146 Hamburg, Germany
[2] Univ Hamburg, Bioctr Grindel, Zool Museum, D-20146 Hamburg, Germany
[3] Royal Belgian Inst Nat Sci, Direct Nat Environm, B-1000 Brussels, Belgium
[4] Forschungsinst, D-60325 Frankfurt, Germany
[5] Na Museum Senckenberg, D-60325 Frankfurt, Germany
[6] Bavarian State Collect Zool, D-81247 Munich, Germany
[7] Biocentrum Grindel, German Ctr Marine Biodivers Res DZMB, D-20146 Hamburg, Germany
[8] Ruhr Univ Bochum, Dept Ecol Evolut & Biodivers, D-44780 Bochum, Germany
[9] Univ Fed Rio Grande do Norte, Lab Geol & Geofis Marinha & Monitoramento Ambient, Programa Posgrad Geodinam & Geofis, Dept Geol,Ctr Ciencias Exatas, BR-59072970 Natal, RN, Brazil