[1] Woods Hole Oceanog Inst, Woods Hole, MA 02543 USA
[2] Univ S Alabama, Dauphin Isl Sea Lab, Dauphin Isl, AL USA
[3] Texas A&M Univ Galveston, Dept Marine & Coastal Environm Sci, Galveston, TX USA
[4] Mississippi State Univ Extens, Mississippi Alabama Sea Grant, Florida Sea Grant, Biloxi, FL USA