[1] Univ Autonoma Barcelona, Inst Neurosci, Bellaterra 08193, Spain
[2] Univ Autonoma Barcelona, Dept Cell Biol Physiol & Immunol, Barcelona, Spain
[3] Inst Salud Carlos III, Ctr Invest Biomed Red Enfermedades Neurodegenerat, Madrid, Spain
[4] Univ Autonoma Barcelona, Dept Biochem & Mol Biol, Barcelona, Spain
[5] Vall DHebron Inst Recerca VHIR, Unitat Mixta UAB VHIR, Barcelona, Spain
[6] Inst Catala Recerca & Estudis Avancats ICREA, Barcelona, Spain