[1] KNAW, Hubrecht Inst, NL-3584 CT Utrecht, Netherlands
[2] UMC Utrecht, NL-3584 CT Utrecht, Netherlands
[3] Univ Groningen, Univ Med Ctr Groningen, Dept Cell Biol, Groningen, Netherlands
[4] Univ Cote Azur, CNRS, Inst Pharmacol Mol & Cellulaire, F-06560 Valbonne, France