[1] Natl Inst Environm Studies, Tsukuba, Ibaraki 3058506, Japan
[2] Beijing Normal Univ, State Key Lab Earth Surface Proc & Resource Ecol, Beijing 100875, Peoples R China
[3] Beijing Normal Univ, Coll Global Change & Earth Syst Sci, Beijing 100875, Peoples R China
[4] Chinese Acad Sci, Inst Tibetan Plateau Res, Key Lab Alpine Ecol & Biodivers, Beijing 100101, Peoples R China