[1] Univ Alaska Fairbanks, Inst Arctic Biol, Dept Biol & Wildlife, EWHALE Lab, 419 Irving 1,902 North Koyukuk Dr, Fairbanks, AK 99775 USA
[2] Michigan State Univ, Dept Entomol, Global Observ Ecosyst Serv, 101 Manly Miles Bldg,1405 South Harrison Rd, E Lansing, MI 48824 USA
[3] US Fish & Wildlife Serv, Kenai Natl Wildlife Refuge, 1 Ski Hill Rd, Soldotna, AK 99669 USA
[4] US Fish & Wildlife Serv, Ecol Serv, 1208-B Main St, Daphne, AL 36526 USA