[1] Univ Algarve, Ctr Marine Sci CCMAR, P-8005139 Faro, Portugal
[2] ISPA Inst Univ, MARE Marine & Environm Sci Ctr, P-1149041 Lisbon, Portugal
[3] Univ Azores, Inst Marine Res, Dept Oceanog & Fisheries, P-9901862 Horta, Portugal
[4] LARSyS, P-1049001 Lisbon, Portugal