[1] Mississippi State Univ, Coastal Res & Extens Ctr, 1815 Popps Ferry Rd, Biloxi, MS 39532 USA
[2] Univ Calif Davis, Ctr Populat Biol, One Shields Ave, Davis, CA 95616 USA
[3] Univ New Brunswick, Dept Biol, POB 4400, Fredericton, NB E3B 5A3, Canada
[4] Univ Vermont, Dept Biol, 63 Carrigan Dr, Burlington, VT 05405 USA