[1] Nagoya Univ, Grad Sch Bioagr Sci, Lab Anim Management & Resources, Aichi 4648601, Japan
[2] Chukyo Womens Univ, Fac Humanities, Aichi 4748651, Japan
[3] Okayama Univ Sci, Dept Biosphere Geoshpere Syst Sci, Okayama 700005, Japan
[4] Nagoya Univ, Grad Sch Bioagr Sci, Lab Anim Management & Resources, Aichi 4648601, Japan