[1] Univ Winnipeg, C FIR, 515 Ave Portage, Winnipeg, MB R3B 2E9, Canada
[2] Univ Manitoba, Dept Bot, Winnipeg, MB R3T 2N2, Canada
[3] Univ Quebec Abitibi Temiscamingue, NSERC UQAT UQAM Ind Res Chair Sustainable Forest, Rouyn Noranda, PQ J9X 5E4, Canada