[1] Chinese Acad Sci, Northeast Inst Geog & Agroecol, Key Lab Wetland Ecol & Environm, Changchun 130102, Jilin, Peoples R China
[2] Landcare Res, 231 Morrin Rd, Auckland, New Zealand
[3] Univ Auckland, Ctr Biodivers & Biosecur, Sch Biol Sci, Auckland, New Zealand