[1] South China Agr Univ, Coll Marine Sci, Guangzhou 510642, Guangdong, Peoples R China
[2] Fisheries & Oceans Canada, St Andrews Biol Stn, St Andrews, NB E5B 2L9, Canada
[3] Univ New Brunswick, Biol Dept, St John, NB E2L 4L5, Canada
[4] Univ New Brunswick, Ctr Coastal Studies & Aquaculture, St John, NB E2L 4L5, Canada