[1] Nanjing Univ Informat Sci & Technol, Sch Marine Sci, Nanjing 210044, Jiangsu, Peoples R China
[2] Guangxi Univ, Sch Marine Sci, Nanning 530004, Peoples R China
[3] Guangxi Univ, Guangxi Lab Study Coral Reefs South China Sea, Nanning 530004, Peoples R China
[4] Univ Hawaii Manoa, Dept Mol Biosci & Bioengn, 1955 East West Rd, Honolulu, HI 96822 USA
[5] Luxembourg Inst Hlth, Dept Populat Hlth, 1A B,Rue Thomas Edison, L-1445 Strassen, Luxembourg
[6] Chinese Acad Sci, State Key Lab Organ Geochem, Guangzhou Inst Geochemi, Guangzhou 510640, Guangdong, Peoples R China