[1] Chinese Acad Sci, South China Bot Garden, Key Lab Vegetat Restorat & Management Degraded Ec, Xingke Rd 723, Guangzhou 510650, Guangdong, Peoples R China
[2] Univ Chinese Acad Sci, Beijing 100049, Peoples R China
[3] Chinese Acad Sci, Xiaoliang Res Stn Trop Coastal Ecosyst, Maoming 525029, Peoples R China
[4] Univ Lancaster, Lancaster Environm Ctr, Lancaster LA1 4YQ, England
[5] Minist Agr, Agroenvironm Protect Inst, Tianjin 300191, Peoples R China
[6] Marine Biol Lab, Woods Hole, MA 02543 USA