[1] Univ Montpellier, EPHE, IRD, Inst Sci Evolut,UMR CNRS 5554, Pl Eugene Bataillon, F-34095 Montpellier 5, France
[2] Museo La Plata, Lab Sistemat & Biol Evolut LASBE, Paseo Bosque S-N,B1900FWA, La Plata, Argentina
[3] Consejo Nacl Invest Cient & Tecn, RA-1033 Buenos Aires, DF, Argentina
[4] Univ Nacl Mayor San Marcos, Museo Hist Nat, Dept Paleontol Vertebrados, Ave Arenales 1256, Lima 11, Peru
[5] Univ Toulouse, CNRS, LMTG, Geosci Environm Toulouse,UPS SVT OMP,IRD, 14 Ave Edouard Belin, F-31400 Toulouse, France
[6] Convenio IRD PeruPetro, Av Luis Aldana 320, Lima, Peru
[7] Univ Paris 06, Sorbonne Univ, CNRS, Museum Natl Hist Nat,MNHN,CR2P, CP 38,8 Rue Buffon, F-75231 Paris 05, France
[8] Museo La Plata, Div Paleontol Vertebrados, Paseo Bosque S-N,B1900FWA, La Plata, Argentina
[9] Univ Oslo, Dept Geosci, POB 1047 Blindem, NO-0316 Oslo, Norway
[10] Case Western Reserve Univ, Sch Med, Dept Anat, Cleveland, OH 44106 USA
[11] Geol Survey Norway, N-7491 Trondheim, Norway
[12] Smithsonian Trop Res Inst, Ctr Trop Paleoecol & Archeol, Balboa 084303092, Ancon Aa, Panama
[13] Goethe Univ Frankfurt, Dept Ecol & Evolut, Max von Laue Str 13, D-60438 Frankfurt, Germany
[14] Paleosedes EU, Tv 27 57-49 Campin, Bogota, Colombia
[15] Consejo Nacl Invest Cient & Tecn, Museo Paleontol Egidio Feruglio, Av Fontana 140, Trelew, Argentina
[16] Field Museum Nat Hist, 1400 Lake Shore Dr, Chicago, IL 60605 USA
[17] CCT CONICET Mendoza, IANIGLA, Ave Ruiz Leal S-N,Parque Gen San Martin, RA-5500 Mendoza, Argentina
[18] Columbia Univ, Lamont Doherty Earth Observ, Dept Earth & Environm Sci, Palisades, NY 10964 USA
[19] Amer Museum Nat Hist, Div Vertebrate Paleontol, New York, NY 10024 USA
[20] Nat Hist Museum Los Angeles Cty, Dept Mammal, Los Angeles, CA USA
[21] Natl Museum Nat Hist, Smithsonian Inst, Dept Paleobiol, Washington, DC 20560 USA
[22] Nat Biodivers Ctr, POB 9517, NL-2300 RA Leiden, Netherlands