[1] E Carolina Univ, Greenville, NC 27858 USA
[2] UNC Coastal Studies Inst, Wanchese, NC 27981 USA
[3] Virginia Inst Marine Sci, Gloucester Point, VA 23062 USA
[4] Univ Washington, Seattle, WA 98195 USA
[5] Natl Inst Water & Atmospher Res, Wellington, New Zealand