[1] Univ British Columbia, Fac Agr Engn, Vancouver, BC V6T 1Z4, Canada
[2] Univ British Columbia, AquaNet, Vancouver, BC V6T 1Z4, Canada
[3] Simon Fraser Univ, Dept Biol Sci, Burnaby, BC V5A 1S6, Canada
[4] Natl Res Council Canada, Inst Marine Biosci, Halifax, NS B3H 4P4, Canada