[1] Fisheries & Oceans Canada, Great Lakes Lab Fisheries & Aquat Sci, Burlington, ON L7S 1A1, Canada
[2] Univ Massachusetts, Dept Environm Conservat, 160 Holdsworth Way, Amherst, MA 01003 USA
[3] Carleton Univ, Dept Biol, Fish Ecol & Conservat Physiol Lab, 1125 Colonel By Dr, Ottawa, ON K1S 5B6, Canada