[1] Univ Penn, Clin Studies Philadelphia, Philadelphia, PA 19104 USA
[2] Waisman Ctr Mental Retardat & Human Dev, Madison, WI USA
[3] McPherson Eye Res Inst, Madison, WI USA
[4] Univ Penn, Dept Biochem, Philadelphia, PA 19104 USA
[5] Univ Wisconsin, Dept Ophthalmol, Madison, WI USA