[1] Burnet Inst, Ctr Biomed Res, Melbourne, Vic, Australia
[2] Monash Univ, Melbourne, Vic 3004, Australia
[3] Boston Univ, Boston, MA 02215 USA
[4] Johns Hopkins Univ, Baltimore, MD USA
[5] Univ Melbourne, Dept Microbiol & Immunol, Melbourne, Vic, Australia