[1] Guizhou Univ, Coll Life Sci, Dept Ecol, Guiyang 550025, Guizhou, Peoples R China
[2] Guizhou Univ, Coll Life Sci, Minist Educ, Key Lab Plant Resource Conservat & Germplasm Inno, Guiyang, Peoples R China
[3] Chinese Acad Sci, Ctr Monsoon Syst Res, Inst Atmospher Phys, Beijing, Peoples R China
[4] Univ Chinese Acad Sci, Coll Earth & Planetary Sci, Beijing, Peoples R China