[1] CSIC, Estn Biol Donana, Integrat Ecol Grp, C Americo Vespucio S-N, Seville 41092, Spain
[2] Univ Zurich, Dept Evolutionary Biol & Environm Studies, Winterthurerstr 190, CH-8057 Zurich, Switzerland
[3] McGill Univ, Dept Biol, 1205 Docteur Penfield, Montreal, PQ H3A 1B1, Canada
[4] Univ Seville, CSIC, Inst Microelect Sevilla, C Americo Vespucio S-N, Seville 41092, Spain