[1] Univ Helsinki, Dept Forest Sci, BoPOB 27, FIN-00014 Helsinki, Finland
[2] Finnish Geospatial Res Inst, Dept Remote Sensing & Photograrranetry, Geodeetinrinne 2, Masala 02431, Finland
[3] Finnish Geospatial Res Inst, Ctr Excellence Laser Scanning Res, Geodeetinrinne 2, Masala 02431, Finland
[4] Univ Eastern Finland, Sch Forest Sci, POB 111, Joensuu 80101, Finland
[5] Univ British Columbia, Dept Forest Resources Management, 2424 Main Mall, Vancouver, BC V6T 1Z4, Canada