[1] Swiss Fed Inst Aquat Sci & Technol, Eawag, Dubendorf, Switzerland
[2] ETH, Swiss Fed Inst Technol, Inst Environm Engn, Zurich, Switzerland
[3] Carnegie Inst Sci, Dept Global Ecol, Stanford, CA USA
[4] ETH, Swiss Fed Inst Technol, Inst Biogeochem & Pollutant Dynam, Zurich, Switzerland