[1] Univ Fed Rio Grande do Norte, Programa Posgrad Sistemat & Evolucao, Natal, RN, Brazil
[2] SUNY Coll Cortland, Dept Biol Sci, Cortland, NY 13045 USA
[3] Inst Nacl Pesquisas Amazonia INPA, Dept Biodiversidade CBio, Manaus, Amazonas, Brazil
[4] Univ Toronto, Dept Nat Hist, Royal Ontario Museum, Toronto, ON, Canada
[5] Univ Toronto, Dept Ecol & Evolutionary Biol, Toronto, ON, Canada
[6] Univ Fed Rio Grande do Norte, Dept Bot & Zool, Natal, RN, Brazil
[7] Univ Fed Paraiba, Programa Posgrad Sistemat & Evolucao, Joao Pessoa, Paraiba, Brazil