[1] CSIRO Marine & Atmospher Res, Hobart, Tas 7001, Australia
[2] Univ Tasmania, Antarctic Wildlife Res Unit, Sch Zool, Hobart, Tas 7001, Australia
[3] Univ St Andrews, NERC Sea Mammal Res Unit, Gatty Marine Lab, St Andrews KY16 8LB, Fife, Scotland
[4] CSIRO Math & Informat Sci, Hobart, Tas 7001, Australia