[1] Michigan State Univ, Dept Biochem & Mol Biol, E Lansing, MI 48824 USA
[2] Michigan State Univ, Ctr Mitochondrial Sci & Med, E Lansing, MI 48824 USA
[3] Univ Tampere, Inst Biomed Technol, FIN-33101 Tampere, Finland
[4] Univ N Carolina, Lineberger Canc Res Ctr, Chapel Hill, NC 27599 USA