[1] Univ Cambridge, Dept Plant Sci, Forest Ecol & Conservat Grp, Cambridge CB2 3EA, England
[2] Univ Reunion, UMR Peuplements Vegetaux & Bioagresseursen Milieu, F-97715 St Denis Messageries, France
[3] Landcare Res, Lincoln 7640, New Zealand
[4] UCL, Dept Geog, London WC1E 6BT, England
[5] Univ Florida, Dept Biol, Gainesville, FL 32611 USA