[1] Univ Western Cape, Dept Biodivers & Conservat Biol, Private Bag X17, ZA-7535 Bellville, South Africa
[2] South African Natl Biodivers Inst, Compton Herbarium, Private Bag X7, Claremont, South Africa
[3] Univ Western Australia, Sch Plant Biol, 35 Stirling Hwy, Perth, WA 6009, Australia
[4] Univ Johannesburg, Dept Bot & Plant Biotechnol, POB 524, ZA-2006 Auckland Pk, South Africa