[1] Univ Montana, WA Franke Coll Forestry & Conservat, Wildlife Biol Program, Missoula, MT 59812 USA
[2] Colorado State Univ, Dept Biol, Ft Collins, CO 80523 USA
[3] Colorado State Univ, Grad Degree Program Ecol, Ft Collins, CO 80523 USA
[4] Michigan State Univ, WK Kellogg Biol Stn, Dept Integrat Biol, Hickory Corners, MI 49060 USA
[5] Michigan State Univ, Ecol Evolutionary Biol & Behav Program, E Lansing, MI 48824 USA
[6] Univ Montana, Div Biol Sci, Missoula, MT 59812 USA
[7] Univ Alaska Southeast, Biol & Marine Biol Program, Juneau, AK USA