[1] Sorbonne Univ, Integrat Biol Marine Models LBI2M, Stn Biol, CNRS,UMR 8227, Pl Georges Teissier, F-29680 Roscoff, France
[2] Sorbonne Univ, Int Res Lab IRL 3614, CNRS,Evolutionary Biol & Ecol Algae,EBEA, Stn Biol,PUC,UACH, F-29680 Roscoff, France
[3] Helmholtz Ctr Ocean Res GEOMAR, Dusternbrooker Weg 20, D-24105 Kiel, Germany
[4] Univ Brest Western Brittany, UBO European Inst Marine Studies IUEM, Lab Sci Environm Marin, IUEM,UMR 6539 LEMAR,Technopole Brest Iroise, Rue Dumt Urville, F-29280 Plouzane, France