[1] ARC, Inst Soil Climate & Water, Belvedere St 600, ZA-0001 Pretoria, South Africa
[2] Univ Pretoria, Dept Plant & Soil Sci, ZA-0028 Pretoria, South Africa
[3] Univ Goettingen, Trop Plant Prod & Agr Syst Modelling TROPAGS, Grisebachstr 6, D-37077 Gottingen, Germany
[4] Univ Goettingen, Ctr Biodivers & Sustainable Land Use CBL, Buesgenweg 1, D-37077 Gottingen, Germany
[5] Univ Free State, Dept Soil Crop & Climate Sci, ZA-9300 Bloemfontein, South Africa
[6] Univ Venda, Dept Zool, ZA-0950 Thohoyandou, South Africa
[7] Swiss Fed Inst Technol, Dept Dev Econ & Cooperat, CH-8093 Zurich, Switzerland