[1] Hellen Ctr Marine Res, Inst Oceanog, GR-71003 Iraklion, Crete, Greece
[2] Univ Crete, Dept Biol, Marine Ecol Lab, GR-71409 Iraklion, Crete, Greece
[3] Univ So Denmark, Inst Biol, DK-5230 Odense M, Denmark
[4] Inst Mediterrani Estudis Avancats CSIC UIB, Dept Global Change, SP-07190 Esporles, Illes Balears, Spain