[1] Natl Chung Hsing Univ, Dept Entomol, 250 Kuo Kuang Rd, Taichung 402, Taiwan
[2] Univ Edinburgh, Inst Evolutionary Biol, Kings Bldg,West Mains Rd, Edinburgh EH9 3JT, Midlothian, Scotland
[3] Univ Szeged, Dept Ecol, Kozep Fasor 52, Szeged, Hungary
[4] Natl Food Chain Safety Off, Directorate Plant Protect Soil Conservat & Agrien, Plant Hlth & Mol Biol Lab, Budaorsi Str 141-145, H-1118 Budapest, Hungary