[1] Natl Marine Fisheries Serv, NE Fisheries Sci Ctr, Woods Hole, MA 02543 USA
[2] Dept Environm Affairs & Tourism, ZA-8012 Cape Town, South Africa
[3] Univ Cape Town, Marine Res MA RE Inst, ZA-7702 Rondebosch, South Africa
[4] CSIC, ICM, Inst Marine Sci, Barcelona 08002, Spain
[5] Inst Rech Dev, UMR EME 212, F-34203 Sete, France
[6] Inst Invest Pescas & Mar IPIMAR INRB, P-1449006 Lisbon, Portugal
[7] Dalhousie Univ, Dept Biol, Halifax, NS B3H 4J1, Canada