[1] Princeton Univ, Dept Ecol & Evolutionary Biol, Princeton, NJ 08544 USA
[2] Farallon Inst, Petaluma, CA 94952 USA
[3] Ctr Biodivers Theory & Modeling, Moulis, France
[4] Stockholm Univ, Stockholm Resilience Ctr, Stockholm, Sweden
[5] Oregon State Univ, Coll Earth Ocean & Atmospher Sci, Corvallis, OR 97331 USA