[1] Shanghai Jiao Tong Univ, Sch Agr & Biol, Shanghai 201101, Peoples R China
[2] Chinese Acad Sci, Inst Subtrop Agr, Key Lab Subtrop Agr Ecol, Changsha 410125, Hunan, Peoples R China
[3] Ehime Univ, United Grad Sch Agr Sci, Matsuyama, Ehime 7908566, Japan
[4] Kochi Univ, Fac Agr, Kochi 7838502, Japan