[1] Univ Tehran, Coll Sci, Sch Biol, Tehran, Iran
[2] Ferdowsi Univ Mashhad, Fac Sci, Dept Biol, Mashhad, Iran
[3] Ferdowsi Univ Mashhad, Fac Sci, Inst Appl Zool, Res Dept Zool Innovat, Mashhad, Iran
[4] RAS, Inst Biol Problems North, Magadan, Russia
[5] Univ Free State, Dept Zool & Entomol, Bloemfontein, South Africa
[6] Univ Furku, Zool Museum, Turku, Finland