[1] Univ Nacl Cordoba, Inst Multidisciplinario Biol Vegetal, RA-5000 Cordoba, Argentina
[2] Consejo Nacl Invest Cient & Tecn, RA-5000 Cordoba, Argentina
[3] Univ Grenoble 1, Lab Ecol Alpine, UMR 5553, CNRS, F-38042 Grenoble 09, France
[4] Univ Grenoble 1, Stn Alpine Joseph Fourier, UMS 2925, CNRS, F-38042 Grenoble 09, France
[5] CNRS, UMR 5175, Ctr Ecol Fonct & Evolut, F-34293 Montpellier 05, France
[6] CNRS MAE, Ctr Francais Rech Sci Soci, Prague 12800 2, Czech Republic
[7] Australian Natl Univ, Natl Ctr Epidemiol & Populat Hlth, Canberra, ACT 0200, Australia