[1] Univ Libre Bruxelles, Lab Physiol & Genet Mol Plantes, Campus Plaine,CP242,Bld Triomphe, B-1050 Brussels, Belgium
[2] Fac Univ Sci Agron Gembloux, Unite Sol Ecol Terr, Lab Geopedol, Gembloux, Belgium
[3] Univ Libre Bruxelles, Lab Ecol Vegetale & Biogeochim, B-1050 Brussels, Belgium
[4] Univ Lubumbashi, Fac Sci Agron, Lubumbashi, DEM REP CONGO